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Task

ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot substitutions, AUC. Scored with AUC on ProteinGym substitution DMS assays. Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

21 evaluations · 21 metric rows

Overview

Zero-shot substitutions, AUC. Scored with AUC on ProteinGym substitution DMS assays. Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Consult the linked sources for architecture or protocol details. Missing evidence is not evidence of a missing capability.

Evaluation design

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Benchmarks

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Recorded evaluations

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Run instructions

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Published comparisons

Explore the results reported under one evaluation protocol. Each figure keeps its source, dataset and metric together; it is not a ranking across studies. The pooled view gathers every source table that reports the same metric and names what it does not hold constant.

ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

auc (fraction) · Higher values are better for this metric.

Every method ProteinGym reports on Zero-shot substitutions, AUC, scored with AUC on ProteinGym substitution DMS assays.

Evaluation protocol · ProteinGym substitution DMS assays (ProteinGym split)

  1. Site-Independent · Configuration · Author-reported evaluation0.696
  2. WaveNet · Configuration · Author-reported evaluation0.707
  3. EVmutation · Configuration · Author-reported evaluation0.716
  4. DeepSequence (ensemble) · Configuration · Author-reported evaluation0.729
  5. EVE (ensemble) · Configuration · Author-reported evaluation0.741
  6. GEMME · Configuration · Author-reported evaluation0.749
  7. UniRep · Configuration · Author-reported evaluation0.605
  8. CARP (640M) · Configuration · Author-reported evaluation0.701
  9. ESM-1b · Configuration · Author-reported evaluation0.719
  10. ESM-2 (15B) · Configuration · Author-reported evaluation0.720
  11. RITA XL · Configuration · Author-reported evaluation0.707
  12. ESM-1v (ensemble) · Configuration · Author-reported evaluation0.723
  13. ProGen2 XL · Configuration · Author-reported evaluation0.717
  14. VESPA · Configuration · Author-reported evaluation0.742
  15. UniRep evotuned · Configuration · Author-reported evaluation0.693
  16. Tranception L · Configuration · Author-reported evaluation0.739
  17. TranceptEVE L · Configuration · Author-reported evaluation0.751
  18. ESM-IF1 · Configuration · Author-reported evaluation0.730
  19. MIF-ST · Configuration · Author-reported evaluation0.718
  20. ProteinMPNN · Configuration · Author-reported evaluation0.639

Source order is preserved. Plotted marks show point estimates; uncertainty, where reported, is retained in the printed values and table. Differences do not establish statistical significance.

ProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, column(Zero-shot substitutions, AUC)
Values, uncertainty and evidence
auc: original source values
Tested entityPrinted valueUncertaintyEvidence
Site-Independent · Configuration0.696 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Alignment- Site-Independent), column(Zero-shot substitutions, AUC)
WaveNet · Configuration0.707 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(based WaveNet), column(Zero-shot substitutions, AUC)
EVmutation · Configuration0.716 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(EVmutation), column(Zero-shot substitutions, AUC)
DeepSequence (ensemble) · Configuration0.729 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(DeepSequence (ensemble)), column(Zero-shot substitutions, AUC)
EVE (ensemble) · Configuration0.741 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(EVE (ensemble)), column(Zero-shot substitutions, AUC)
GEMME · Configuration0.749 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(GEMME), column(Zero-shot substitutions, AUC)
UniRep · Configuration0.605 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Protein UniRep), column(Zero-shot substitutions, AUC)
CARP (640M) · Configuration0.701 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(language CARP (640M)), column(Zero-shot substitutions, AUC)
ESM-1b · Configuration0.719 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ESM-1b), column(Zero-shot substitutions, AUC)
ESM-2 (15B) · Configuration0.720 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ESM-2 (15B)), column(Zero-shot substitutions, AUC)
RITA XL · Configuration0.707 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(RITA XL), column(Zero-shot substitutions, AUC)
ESM-1v (ensemble) · Configuration0.723 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ESM-1v (ensemble)), column(Zero-shot substitutions, AUC)
ProGen2 XL · Configuration0.717 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ProGen2 XL), column(Zero-shot substitutions, AUC)
VESPA · Configuration0.742 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(VESPA), column(Zero-shot substitutions, AUC)
UniRep evotuned · Configuration0.693 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Hybrid UniRep evotuned), column(Zero-shot substitutions, AUC)
MSA Transformer (ensemble) · Configuration0.738 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(MSA Transformer (ensemble)), column(Zero-shot substitutions, AUC)
Tranception L · Configuration0.739 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Tranception L), column(Zero-shot substitutions, AUC)
TranceptEVE L · Configuration0.751 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(TranceptEVE L), column(Zero-shot substitutions, AUC)
ESM-IF1 · Configuration0.730 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Inverse ESM-IF1), column(Zero-shot substitutions, AUC)
MIF-ST · Configuration0.718 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Folding MIF-ST), column(Zero-shot substitutions, AUC)
ProteinMPNN · Configuration0.639 fractionNot reportedAuthor-reported evaluation · source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ProteinMPNN), column(Zero-shot substitutions, AUC)
Scope and limitations
  • Author-reported numbers, source checked but not independently reproduced.
  • MSE is better when lower; every other metric here is better when higher.
  • Zero-shot, supervised and indel benchmarks use different protocols and assay sets, so their figures are not comparable to each other.

Source transcription and grouping reviewed by automated source review on 2026-09-18. These experiments were not independently reproduced by rewire.

Tested entities and results

Release 2026-09-17-134cd1815de8 · 21 evaluations · 21 metric rows. Different protocols are not a single leaderboard. Where several source tables report the same metric, the published comparisons above offer a pooled view that names what it does not hold constant.

Results grouped by the exact reported evaluation
Metric and findingCoverage and uncertaintyEvidence
CARP (640M) on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.701 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(language CARP (640M)), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

DeepSequence (ensemble) on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.729 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(DeepSequence (ensemble)), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

ESM-1b on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.719 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ESM-1b), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

ESM-1v (ensemble) on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.723 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ESM-1v (ensemble)), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

ESM-2 (15B) on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.720 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ESM-2 (15B)), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

ESM-IF1 on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.730 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Inverse ESM-IF1), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

EVE (ensemble) on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.741 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(EVE (ensemble)), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

EVmutation on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.716 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(EVmutation), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

GEMME on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.749 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(GEMME), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

MIF-ST on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.718 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Folding MIF-ST), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

MSA Transformer (ensemble) on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.738 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(MSA Transformer (ensemble)), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

ProGen2 XL on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.717 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ProGen2 XL), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

ProteinMPNN on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.639 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(ProteinMPNN), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

RITA XL on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.707 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(RITA XL), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

Site-Independent on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.696 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Alignment- Site-Independent), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

TranceptEVE L on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.751 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(TranceptEVE L), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

Tranception L on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.739 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Tranception L), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

UniRep evotuned on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.693 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Hybrid UniRep evotuned), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

UniRep on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.605 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(Protein UniRep), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

VESPA on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.742 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(VESPA), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

WaveNet on ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC

Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.

Author-reported evaluation · Evaluation metadata: source checked

0.707 auc

Unit: fraction · Direction: higher

Uncertainty: Not reported

Scored: Not reported · Eligible: Not reported

source checkedProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design · Table 2, row(based WaveNet), column(Zero-shot substitutions, AUC)

Source checking is not independent reproduction.

Evidence table

Inspect claims, sources and review details

Trace each statement to its source and review. A context-only reference supports the record generally; it does not verify an individual field. Source checking does not reproduce an experiment.

One row per statement and cited source. Multiple citations are not independent evaluations. Shared locators are labelled explicitly.

1 evidence row matching the loaded filters

Claims, original sources and review scope · Release 2026-09-17-134cd1815de8
Property and statementOriginal source and locationReview and provenance
Relationship: part of

discovery-benchmark-proteingym

Individual claims
ProteinGym: Large-Scale Benchmarks for Protein Fitness Prediction and Design

Original source ↗

Table 2, column(Zero-shot substitutions, AUC)

Version: Primary full-text snapshot retrieved 2026-09-17; exact bytes pinned by SHA-256
Retrieved: 2026-09-17T08:06:28.522924+00:00

source checked

automated source review · 2026-09-18

Audit details

Primary-source transcription with no human sign-off and no independent reproduction.

Field: links:part_of:discovery-benchmark-proteingym

Claim: proteingym-association-zs-sub-auc

Source artifact SHA-256: a3b08cc4a6befd64620cf0f287d78d55a36dc2639955f52c5655f83833c50104

Hash scope: Exact retrieved primary paper artifact bytes.

Inspected artifact

Sources and history

Release 2026-09-17-134cd1815de8 · Record review: source checked

1 source records and release historyDownload this release
Technical metadata and extraction receipts

Stable ID: proteingym-task-zs-sub-auc

areas
proteins-complexes
tasks
Zero-shot substitutions, AUC
metric
AUC
metric direction
higher
dataset
ProteinGym substitution DMS assays
protocol
Zero-shot scoring of substitution assays, averaged over assays with the correction the paper describes.
source locator
Table 2, column(Zero-shot substitutions, AUC)
comparison panels
id: proteingym-panel-zs-sub-auc; title: ProteinGym ZS-SUB-AUC: Zero-shot substitutions, AUC; protocol id: proteingym-task-zs-sub-auc; dataset id: proteingym-dataset-proteingym-substitution-dms-assays; metric: auc; unit: fraction; direction: higher; result ids: proteingym-result-site-independent-zs-sub-auc-auc; proteingym-result-wavenet-zs-sub-auc-auc; proteingym-result-evmutation-zs-sub-auc-auc; proteingym-result-deepsequence-ensemble-zs-sub-auc-auc; proteingym-result-eve-ensemble-zs-sub-auc-auc; proteingym-result-gemme-zs-sub-auc-auc; proteingym-result-unirep-zs-sub-auc-auc; proteingym-result-carp-640m-zs-sub-auc-auc; proteingym-result-esm-1b-zs-sub-auc-auc; proteingym-result-esm-2-15b-zs-sub-auc-auc; proteingym-result-rita-xl-zs-sub-auc-auc; proteingym-result-esm-1v-ensemble-zs-sub-auc-auc; proteingym-result-progen2-xl-zs-sub-auc-auc; proteingym-result-vespa-zs-sub-auc-auc; proteingym-result-unirep-evotuned-zs-sub-auc-auc; proteingym-result-msa-transformer-ensemble-zs-sub-auc-auc; proteingym-result-tranception-l-zs-sub-auc-auc; proteingym-result-trancepteve-l-zs-sub-auc-auc; proteingym-result-esm-if1-zs-sub-auc-auc; proteingym-result-mif-st-zs-sub-auc-auc; proteingym-result-proteinmpnn-zs-sub-auc-auc; source ids: evidence-expansion-proteingym-a3b08cc4; source locator: Table 2, column(Zero-shot substitutions, AUC); context: Every method ProteinGym reports on Zero-shot substitutions, AUC, scored with AUC on ProteinGym substitution DMS assays.; caveats: Author-reported numbers, source checked but not independently reproduced.; MSE is better when lower; every other metric here is better when higher.; Zero-shot, supervised and indel benchmarks use different protocols and assay sets, so their figures are not comparable to each other.; review: method: automated_source_review; date: 2026-09-18
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